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New material of Early Pleistocene Sus lydekkeri (Artiodactyla, Mammalia) from Luotuo Hill in Dalian, Northeast China

DONG Wei LIU Wen-Hui2  LIU Si-Zhao3  WANG Yuan1  LIU Jin-Yuan3  JIN Chang-Zhu1   

  1. (1 Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing 100044   dongwei@ivpp.ac.cn)
    (2 National Museum of China Beijing 100006)
    (3 Dalian Natural History Museum Dalian 116023)

辽宁大连骆驼山的早更新世李氏野猪新材料

董  为1   刘文晖2   刘思昭3   王  元1   刘金远3   金昌柱1   

  1. (1 中国科学院古脊椎动物与古人类研究所,脊椎动物演化与人类起源重点实验室 北京 100044)
    (2 中国国家博物馆环境考古研究所 北京 100006)
    (3 大连自然博物馆 大连 116023)

Abstract:

Chronologically ranged from the Early to Middle Pleistocene, Sus lydekkeri is widely seen in mammalian faunas of the Palearctic and transitional zone of China. More than 20 localities with its remains were reported, but most specimens collected were fragmental and only two nearly complete skulls were described, one from the Middle Pleistocene Peking Man Site at Zhoukoudian in Beijing and another from the Early Pleistocene Yangshuizhan in the Nihewan Basin. The present paper described a third nearly complete and the best-preserved skull of S. lydekkeri so far and other specimens from Jinyuan Cave deposits (paleomagnetically dated 1.1–1.8 Ma) in Dalian, northeastern China. The comparisons of the specimens of S. lydekkeri from different localities show that the intraspecific variation of S. lydekkeri is large and it overlaps with that of S. peii, S. strozzii and S. scrofa in considerable scales. Based on the comparison with available specimens, the most significant difference between S. lydekkeri and S. peii is that the snout of the former is moderately longer than the latter. While the morphological differences between S. lydekkeri and S. strozzii are not very significant. Whereas those between S. lydekkeri and S. scrofa are mostly shown on the canine apparatus, more developed in the former than in the latter. The canine eminence in Sus is a derived character from its Miocene ancestor, evolved into its maximum degree in the Early Pleistocene and reduced gradually in the Late Pleistocene. And that of Dalian skull is the most robust representative among the available specimens of S. lydekkeri. Although the materials are enriched gradually, more complete specimens and larger samples are needed to figure out the clear images of Sus species in face of their puzzling intraspecific variations.

Key words: Luotuo Hill in Dalian, Early Pleistocene, Sus lydekkeri, morphology, intraspecific variation

摘要:

李氏野猪(Sus lydekkeri)是分布于中国古北界和过渡带的早更新世和中更新世哺乳动物群中的常见成员。虽然迄今报道过的产李氏野猪的化石地点逾20个,但是大部分地点产出的标本均比较破碎,或为单独的牙齿。此前仅在周口店的中更新世北京人遗址和泥河湾盆地的早更新世扬水站地点各发现一具近于完整的头骨。记述了产于大连金远洞早更新世堆积(古地磁年龄为1.1–1.8 Ma)的李氏野猪头骨及其他标本。这是第3件近于完整也是目前保存最好的头骨。与其他地点产出的李氏野猪相比可见,李氏野猪的种内变异较大,并与裴氏猪(S. peii)、斯氏猪(S. strozzii)和野猪(S. scrofa)的种内变异范围有较大的重叠。与现有标本的比较显示,李氏野猪和裴氏猪的主要差异在于李氏野猪的鼻吻部比裴氏猪略长。而李氏野猪与斯氏猪的形态差别并不显著,与野猪的差别主要体现在李氏野猪的犬齿组件比野猪的发育。猪属的犬齿盖是从其中新世祖先那里衍生下来的一个特征,在早更新世时演化到顶峰,然后在晚更新世逐渐退化。而大连的李氏野猪头骨上的犬齿盖在已知标本中是最发育的。虽然野猪的材料在不断积累,但是要搞清猪属各个种的清晰面貌及其之间的关系尚需找到更完整的标本并积累更大的样本。

关键词: 大连骆驼山, 早更新世, 李氏野猪, 形态学, 种内变异