古脊椎动物学报 ›› 2021, Vol. 59 ›› Issue (1): 45-63.DOI: 10.19615/j.cnki.1000-3118.200821
张立民1,2,3, 董为1,2, 倪喜军1,2,3,*(), 李强1,2,3,*()
收稿日期:
2020-07-02
出版日期:
2021-01-20
发布日期:
2021-01-20
通讯作者:
*nixijun@ivpp.ac.cn;基金资助:
ZHANG Li-Min1,2,3, DONG Wei1,2, NI Xi-Jun1,2,3,*(), LI Qiang1,2,3,*()
Received:
2020-07-02
Published:
2021-01-20
Online:
2021-01-20
摘要:
近百年来,在内蒙古中部地区发现了丰富的新近纪哺乳动物化石,命名和描述了10余个哺乳动物群,其时代跨越早中新世至早上新世。它们已经成为建立中国北方新近纪哺乳动物生物年代地层框架的重要依据。早在1959年中苏古生物联合考察过程中就已经发现土城子地区有丰富的哺乳动物化石,但这些化石以大-中型哺乳动物为主,缺乏小哺乳动物。土城子动物群的构成和其在中国北方新近纪哺乳动物年代地层框架中的位置长期以来不是特别明确。报道了在土城子新发现的6种小哺乳动物化石。根据Lophocricetus grabaui-Sinocricetus zdanskyi-Prosiphneus licenti-Hansdebruijnia pusilla-Moschus grandaevus的组合特点,认为土城子动物群的年代应该可以很好地约束在晚中新世(或者保德期)的晚期。土城子动物群明显比内蒙古的宝格达乌拉动物群要进步一些,与二登图动物群非常相似,时代上晚于前者,略早于后者。依据动物群整体面貌,推测在晚中新世晚期土城子地区存在森林和草原混合的环境。
中图分类号:
张立民, 董为, 倪喜军, 李强. 晚中新世晚期土城子小哺乳动物组合及土城子动物群在内蒙古中部地区新近纪哺乳动物群序列中的位置. 古脊椎动物学报, 2021, 59(1): 45-63.
ZHANG Li-Min, DONG Wei, NI Xi-Jun, LI Qiang. Late Miocene micromammalian assemblage of Tuchengzi and its biochronological position in Neogene faunal sequence in central Nei Mongol, China. Vertebrata Palasiatica, 2021, 59(1): 45-63.
Fig. 1 Fresh profile of Chilotherium pit in “Dragon bone” hill near Tuchengzi village, Huade Photo taken by LIU Wenhui in 2015; L. abbreviation of layer
Fig. 2 Humeri and mandible of ?Yanshuella primaeva from Tuchengzi, Huade A. IVPP V 26813.3, right distal of humerus; B. V 26813.2, left humerus; C. V 26813.1, right mandible with p3 and m1-3. A1 and B1, A2 and B2, A3 and B3 are in posterior, distal and anterior views, respectively; C1, C2, C3 are in buccal, lingual and occlusal views, respectively. Scale bars equal 1 mm
Tooth | Number | Length | Width | |||
---|---|---|---|---|---|---|
Range | Mean | Range | Mean | |||
M1 | 2 | 3.10-3.20 | 3.15 | 2.26-2.30 | 2.28 | |
M2 | 2 | 2.60-2.65 | 2.63 | 2.15-2.20 | 2.18 | |
M3 | 2 | 2.10-2.15 | 2.13 | 1.95-1.95 | 1.95 | |
m1 | 3 | 2.90-3.35 | 3.10 | 1.65-2.15 | 1.92 | |
m2 | 3 | 2.75-3.10 | 2.88 | 2.20-2.45 | 2.35 | |
m3 | 1 | - | 2.55 | - | 2.15 |
Table 1 Molar measurements of Prosiphneus licenti from Tuchengzi, Huade, Nei Mongol (mm)
Tooth | Number | Length | Width | |||
---|---|---|---|---|---|---|
Range | Mean | Range | Mean | |||
M1 | 2 | 3.10-3.20 | 3.15 | 2.26-2.30 | 2.28 | |
M2 | 2 | 2.60-2.65 | 2.63 | 2.15-2.20 | 2.18 | |
M3 | 2 | 2.10-2.15 | 2.13 | 1.95-1.95 | 1.95 | |
m1 | 3 | 2.90-3.35 | 3.10 | 1.65-2.15 | 1.92 | |
m2 | 3 | 2.75-3.10 | 2.88 | 2.20-2.45 | 2.35 | |
m3 | 1 | - | 2.55 | - | 2.15 |
Tooth | Inventory number | Enamel parameters of upper molars | ||||
---|---|---|---|---|---|---|
(IVPP) | A | B | C | D | ||
M1 | V 26816.1 | 0.1 | 0.3 | 0.6 | 0.6 | |
M1 | V 26816.2 | 0 | 0.55 | 0.55 | 0.6 | |
M2 | V 26816.1 | ? | ? | 0.2 | 0 | |
M2 | V 26816.2 | 0 | 0.25 | 0.25 | 0 | |
M3 | V 26816.1 | 0 | 0.05 | 0.15 | 0 | |
M3 | V 26816.2 | 0.1 | 0.05 | 0.2 | 0 | |
Enamel parameters of lower molars | ||||||
a | b | c | d | e | ||
m1 | V 26816.6 | 0 | 0.25 | 0.2 | 0.3 | 0 |
m1 | V 26816.7 | 0 | 0.45 | 0.25 | 0.4 | 0 |
m2 | V 26816.8 | 0.15 | 0.5 | 0.3 | 0.25 | 0 |
m3 | V 26816.10 | 0.05 | 0.15 | 0.25 | 0.1 | 0 |
Table 2 Molars’ enamel parameters of Prosiphneus licenti from Tuchengzi, Huade, Nei Mongol (mm)
Tooth | Inventory number | Enamel parameters of upper molars | ||||
---|---|---|---|---|---|---|
(IVPP) | A | B | C | D | ||
M1 | V 26816.1 | 0.1 | 0.3 | 0.6 | 0.6 | |
M1 | V 26816.2 | 0 | 0.55 | 0.55 | 0.6 | |
M2 | V 26816.1 | ? | ? | 0.2 | 0 | |
M2 | V 26816.2 | 0 | 0.25 | 0.25 | 0 | |
M3 | V 26816.1 | 0 | 0.05 | 0.15 | 0 | |
M3 | V 26816.2 | 0.1 | 0.05 | 0.2 | 0 | |
Enamel parameters of lower molars | ||||||
a | b | c | d | e | ||
m1 | V 26816.6 | 0 | 0.25 | 0.2 | 0.3 | 0 |
m1 | V 26816.7 | 0 | 0.45 | 0.25 | 0.4 | 0 |
m2 | V 26816.8 | 0.15 | 0.5 | 0.3 | 0.25 | 0 |
m3 | V 26816.10 | 0.05 | 0.15 | 0.25 | 0.1 | 0 |
Fig. 3 Rodent and lagomorph remains from Tuchengzi, Huade A. Sinocricetus zdanskyi, left maxilla with M1-2, IVPP V 26815.1; B. Hansdebruijnia pusilla, left maxilla with M1, V 21817; C. Lophocricetus grabaui, right maxilla with M1-2, V 26814.2; D-G. Prosiphneus licenti: D. left maxilla with M1-3, V 26816.1, E. left m1, V 26816.6, F. left m2, V 26816.8; G. left m3, V 26816.10; H-Q. Ochotona lagreli: H. left mandible with p3-m1, V 21818.1, I. right P2, V 21818.2, J. left P3, V 21818.3, K. right P3, V 21818.6, L. left P4/M1, V 21818.8, M. right M2, V 21818.17, N. right p3, V 21818.19, O. right p3, V 21818.20, P. left p3, V 21818.18, Q. left p4/m1/m2, V 21818.21 A-G, H2, I-Q in occlusal views, H1 in buccal view, H3 in lingual view. Scale bars equal 1 mm
Tooth | Number | Length | Width | |||
---|---|---|---|---|---|---|
Range | Mean | Range | Mean | |||
P2 | 1 | - | 1.0 | - | 2.0 | |
P3 | 4 | 1.35-1.50 | 1.41 | 3.05-3.40 | 3.21 | |
P4/M1 | 7 | 1.40-1.75 | 1.50 | 2.85-3.70 | 3.13 | |
M2 | 3 | 1.75-1.90 | 1.83 | 2.70-3.15 | 3.0 | |
p3 | 4 | 2.0-2.30 | 2.21 | 1.80-2.50 | 2.19 | |
p4/m1/m2 | 4 | 1.35-2.30 | 1.84 | 1.15-2.50 | 1.95 |
Table 3 Tooth measurements of Ochotona lagreli from Tuchengzi, Huade, Nei Mongol (mm)
Tooth | Number | Length | Width | |||
---|---|---|---|---|---|---|
Range | Mean | Range | Mean | |||
P2 | 1 | - | 1.0 | - | 2.0 | |
P3 | 4 | 1.35-1.50 | 1.41 | 3.05-3.40 | 3.21 | |
P4/M1 | 7 | 1.40-1.75 | 1.50 | 2.85-3.70 | 3.13 | |
M2 | 3 | 1.75-1.90 | 1.83 | 2.70-3.15 | 3.0 | |
p3 | 4 | 2.0-2.30 | 2.21 | 1.80-2.50 | 2.19 | |
p4/m1/m2 | 4 | 1.35-2.30 | 1.84 | 1.15-2.50 | 1.95 |
TCZ | ETT | BGD | MH | WLH | |
---|---|---|---|---|---|
Insectivora | |||||
Talpidae | |||||
Yanshuella | primaeva? | primaeva | |||
Rodentia | |||||
Zapodidae | |||||
Lophocricetus | grabaui | grabaui | xianensis | cf. grabaui | |
Cricetidae | |||||
Sinocricetus | zdanskyi | zdanskyi | zdanskyi | ||
Myospalacidae | |||||
Prosiphneus | licenti | eriksoni | eriksoni | murinus | |
Muridae | |||||
Hansdebruijnia | pusilla | pusilla | perpusilla & pusilla | ||
Lagomorpha | |||||
Ochotonidae | |||||
Ochotona | lagreli | lagreli & minor | sp. | lagreli | |
Perissodactyla | |||||
Equidae | |||||
Sinohippus | robustus | sp. | robustus | ||
Hipparion | fossatum | richthofeni | tchikoicum | sp. | teilhardi & platyodus |
Rhinocerotidae | |||||
Chilotherium | gracile | anderssoni | |||
Aceratherium | huadeensis | ||||
Artiodactyla | |||||
Cervidae | |||||
Eostyloceros | blainvillei & triangularis | ||||
Euprox | sp. | ||||
Cervavitus | novorossiae & huadeensis & shanxius | sp. | novorossiae | ||
Moschidae | |||||
Moschus | grandaevus | grandaevus | sp. | ||
Giraffidae | |||||
Palaeotragus | sp. | sp. | sp. | microdon | |
Samotherium | sp. | sinense | |||
Bovidae | |||||
Dorcadoryx | sp. | ||||
Plesiaddax | sp. | depereti |
Table 4 Composition of the Tuchengzi Fauna and a comparison with other related faunas
TCZ | ETT | BGD | MH | WLH | |
---|---|---|---|---|---|
Insectivora | |||||
Talpidae | |||||
Yanshuella | primaeva? | primaeva | |||
Rodentia | |||||
Zapodidae | |||||
Lophocricetus | grabaui | grabaui | xianensis | cf. grabaui | |
Cricetidae | |||||
Sinocricetus | zdanskyi | zdanskyi | zdanskyi | ||
Myospalacidae | |||||
Prosiphneus | licenti | eriksoni | eriksoni | murinus | |
Muridae | |||||
Hansdebruijnia | pusilla | pusilla | perpusilla & pusilla | ||
Lagomorpha | |||||
Ochotonidae | |||||
Ochotona | lagreli | lagreli & minor | sp. | lagreli | |
Perissodactyla | |||||
Equidae | |||||
Sinohippus | robustus | sp. | robustus | ||
Hipparion | fossatum | richthofeni | tchikoicum | sp. | teilhardi & platyodus |
Rhinocerotidae | |||||
Chilotherium | gracile | anderssoni | |||
Aceratherium | huadeensis | ||||
Artiodactyla | |||||
Cervidae | |||||
Eostyloceros | blainvillei & triangularis | ||||
Euprox | sp. | ||||
Cervavitus | novorossiae & huadeensis & shanxius | sp. | novorossiae | ||
Moschidae | |||||
Moschus | grandaevus | grandaevus | sp. | ||
Giraffidae | |||||
Palaeotragus | sp. | sp. | sp. | microdon | |
Samotherium | sp. | sinense | |||
Bovidae | |||||
Dorcadoryx | sp. | ||||
Plesiaddax | sp. | depereti |
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