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New fossils of small and medium-sized bovids from the Early Site of Shanshenmiaozui in Nihewan Basin, North China
TONG Hao-Wen, ZHANG Bei, CHEN Xi, WANG Xiao-Min
Vertebrata Palasiatica    2022, 60 (2): 134-168.   DOI: 10.19615/j.cnki.2096-9899.220413
Abstract   (564 HTML128 PDF(pc) (11536KB)(532)  

Shanshenmiaozui site in Nihewan Basin in North China is a recently discovered Early Pleistocene site which yields rich and diverse mammalian fossils. In the fauna, the small and medium-sized bovid fossils are well represented and can be referred to the following taxa: Spirocerus wongi, Gazella sinensis, Ovis shantungensis and Megalovis piveteaui respectively, among which G. sinensis is the dominate species. S. wongi and G. sinensis are mainly represented by horn-cores and partial skull bones as well as mandibles; in addition, metacarpal and/or metatarsal bones were also recognized for all of the four species. The horn-cores are easy to be identified to the species level, while the dentitions and the postcranial bones underwent a series of examinations and comparisons before getting properly determined and referred to the most approximate taxa. Among the postcranial bones, the metapodials, especially to the metacarpal bones special attentions were paid, which are crucial not only for taxonomic identification, but also for phylogenetic and paleoecological reconstructions; the previously misidentified metapodial specimens in Nihewan fauna were reconsidered in this paper. In the SSMZ fauna, the bovid guild is dominated by Gazella and Bison, which indicates steppe was the most important biome in Nihewan Basin during Early Pleistocene.


Fig. 11 Length vs distal width of metacarpal bones of diverse bovids
The data un-included in Table 4 are from Colbert and Hooijer, 1953; Scott, 1985
Extracts from the Article
The bovid metacarpal bones unearthed at SSMZ site are quite diversified, and most of them are easy to be identified according to their size and proportion, whereas some of them are very similar to each other both in size and stoutness. Therefore, an index should be employed, that’s the robusticity index or stoutness index. Concerning the robusticity index, there exist two calculation methods: one is the relative dimensions of mediolateral diaphysis width and maximum length (SD/GL) ×100) (Maniakas and Kostopoulos, 2017), the other is the distal width/total length (Sher, 1997). Because it’s easier and more practical to measure the distal width, this paper follows Sher’s (1997) method to calculate the robusticity index, i.e. (distal width/greatest length) ×100. Based on the morphological observations and robusticity index calculations, the metacarpal bones of bovids from SSMZ site are quite clearly distinguished (Table 4; Fig. 11).
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