泥河湾山神庙咀早更新世李氏野猪新材料
收稿日期: 2025-01-02
网络出版日期: 2025-07-11
基金资助
国家自然科学基金(42172021);国家自然科学基金(41572003)
版权
New fossil materials of Sus lydekkeri from the Early Pleistocene Shanshenmiaozui site in Nihewan Basin of North China
Received date: 2025-01-02
Online published: 2025-07-11
Copyright
猪科动物是中国更新世哺乳动物群中最常见分子之一,而其系统发育分类和进化关系还远不清楚。在华北泥河湾盆地早更新世山神庙咀遗址新发现的猪科化石材料为了解华北地区猪类的演化提供了新证据。新材料包括雄性个体的部分头骨和下颌骨,它们的大部分牙齿都保存完好。这些标本可归入李氏野猪(Sus lydekkeri), 该种是早更新世华北地区唯一的猪科动物物种。之前报道过的更新世沁县河猪(Potamochoerus chinhsienensis), 其化石材料与河猪的特征相去甚远,依据大小和形态特征暂可将其归入李氏野猪,至少不应当再被视为河猪。李氏野猪与现生野猪(S. scrofa)非常接近,甚至被一些作者视为后者的一个时间亚种,但李氏野猪确实个体明显较大、眶前窝发育、眶下孔位置靠后、雄性下犬齿横截面呈爪哇疣猪型(至少是过渡类型)及颊齿冠面结构简单等,上述特征支持了 “李氏野猪” 作为独立物种的有效性,尤其是早更新世类型的特征更是体型较大,颊齿L/W比值较低,上下M3冠面结构都不超过三个叶,雄性下犬齿完全属于爪哇疣猪型。猪科化石是指示古环境和古气候的重要指标。中国北方和南方更新世动物群中都有大量猪科动物,但南方的猪类动物具有较高的属种多样性,化石发现也更加丰富,这些现象究竟是由不同的起源还是由不同环境所而造成,目前尚不完全清楚。
同号文 , 陈曦 , 张贝 , 孙吉嘉 . 泥河湾山神庙咀早更新世李氏野猪新材料[J]. 古脊椎动物学报, 2025 , 63(3) : 210 -241 . DOI: 10.19615/j.cnki.2096-9899.250711
Suid is one of the most common taxa among the Pleistocene mammalian fauna in China. However, its phylogenetic taxonomy and evolution are far from clear. The newly recovered suid fossil materials from the Early Pleistocene Shanshenmiaozui site in Nihewan Basin in North China provide new insight into the evolution of its kind in North China. The new materials include partial skulls and mandibles of males, and most of their teeth were preserved in situ. The specimens can be referred to the species Sus lydekkeri, which is the only species of Sus found in North China from the Early Pleistocene. The fossils of the once reported Pleistocene species “Potamochoerus chinhsienensis” should also be provisionally included in the species S. lydekkeri according to its form and size, which at least now excludes it from Potamochoerus. The species S. lydekkeri is very close to the extant species S. scrofa, and even was combined into the latter as a chronological subspecies by some authors, while its larger size, prominent preorbital fossa, backwardly positioned infraorbital foramen, frequently appearing verrucosic type (at least the intermediate type of the male lower canine), and simple crown structures support its validness as an independent species. The Early Pleistocene forms are characterized by a larger size, lower L/W ratio of cheekteeth, both upper and lower M3s having no more than three lobes, and the male lower canine is exclusively of a verrucosic type. Suid fossils are a very good indicator of paleoenvironments and paleoclimate. Both the northern and the southern faunas in China contain suid fossils throughout the Pleistocene Epoch, while the southern fauna has prominently more diversified taxa and more abundant fossil collections; whether that is caused by different origins or by environmental modifications is still not clear.
| [1] | Albarella U, Keith M D, Peter A R-C, 2009. Size and shape of the Eurasian wild boar (Sus scrofa), with a view to the reconstruction of its Holocene history. Environ Archaeol, 14: 103-136 |
| [2] | Anezaki T, Hongo H, Kurosawa Y, 2013. Size and shape variation of mandible and mandibular teeth in the Ryukyu wild boar (Sus scrofa riukiuanus). Quat Res, 52(6): 255-264 |
| [3] | Arribas A, Garrido G, 2008. Un nuevo jabalí del género Potamochoerus (Suidae, Artiodactyla, Mammalia) en el plioceno superior terminal euroasiático (Fonelas p-1, Cuenca de guadix, Granada). Cuad Mus Geomin, 10: 337-364 |
| [4] | Azzaroli A, 1975. Remarks on the Pliocene Suidae of Europe. Z S?ugetierkd, 40: 355-367 |
| [5] | Berdondini E, 1992. Suids of the early Villafranchian of Villafranca d'Asti and China. Rend Lincei, 9: 109-124 |
| [6] | Benjamin N R, Crooijmans R P M A, Jordan L R, et al. 2023. Swine global genomic resources: insights into wild and domesticated populations. Mamm Genome, 34: 520-530 |
| [7] | Brands S J comp, 1989-present. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Zwaag, the Netherlands, http://taxonomicon.taxonomy.nl/ |
| [8] | Chen G F, 2004. Artiodactyla. In: Zheng S ed.ed. Jianshi Hominid Site. Beijing: Science Press. 254-308 |
| [9] | Chen X, Wu S M, Wang P, et al. 2020. A report on the Late Pleistocene vertebrate fossils from the Zhangshan locality, Suqian, Jiangsu Province. Acta Anthropol Sin, 39(2): 319-331 |
| [10] | Cherin M, Sorbelli L, Crotti M, et al. 2018. New material of Sus strozzii (Suidae, Mammalia) from the Early Pleistocene of Italy and a phylogenetic analysis of suines. Quat Sci Rev, 194: 94-115 |
| [11] | Chia L P, Wang C, 1978. Hsihoutu—a Culture Site of Early Pleistocene in Shansi Province. Beijing: Cultural Relics Publishing House. 1-85 |
| [12] | Chow M C, 1964. Mammals of “Lantian Man” locality at Lantian, Shensi. Vert PalAsiat, 8(3): 301-308 |
| [13] | Colbert E H, 1935. Siwalik mammals in the American Museum of Natural History. Trans Am Philos Soc, 26: 1-401 |
| [14] | Colbert E H, Hooijer D A, 1953. Pleistocene mammals from the limestone fissures of Szechuan, China. Bull Am Mus Nat Hist, 102: 1-134 |
| [15] | Deng T, Qiu Z X, Wang B Y, et al. 2013. Late Cenozoic biostratigraphy of the Linxia Basin, northwestern China. In: Wang X M, Flynn L J, Fortelius M eds. Fossil Mammals of Asia: Neogene Biostratigraphy and Chronology. New York: Columbia University Press. 243-273 |
| [16] | Dong W, 1999. The Artiodactyla from Hulu cave, Tangshan, Nanjing and the environment of Nanjing man. Acta Anthropol Sin, 18: 270-281 |
| [17] | Dong W, 2008. Early Pleistocene suid (mammal) from the Dajushan, Huainan, Anhui Province (China). Vert PalAsiat, 46(3): 233-246 |
| [18] | Dong W, 2019. Brief history of pigs. Fossils, (1): 2-7 |
| [19] | Dong W, Fang Y S, 2008. New suid materials from the Early Pleistocene of the Tuozi Cave at Tangshan, Jiangsu Province, China. In: Dong W ed. Proceedings of the Eleventh Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press. 53-63 |
| [20] | Dong W, Li Z Y, 2008. Late Pleistocene Artiodactyla (Mammalia) from the Lingjing Site, Xuchang, Henan Province (China). Vert PalAsiat, 46(1): 31-50 |
| [21] | Dong W, Zhang L M, 2014. New materials of Hippopotamodon (Artiodactyla, Mammalia) from southern China. Vert PalAsiat, 16(4): 201-216 |
| [22] | Dong W, Jin C Z, Zheng L T, et al. 2006. Artiodactyla from the Jinpendong site at Wuhu, Anhui Province. Acta Anthropol Sin, 25: 161-171 |
| [23] | Dong W, Fang Y S, Zhang Z H, 2007. Artiodactyla. In: Nanjing Museum, Institute of Archaeology, Jiangsu Province ed. The Early Pleistocene Mammalian Fauna at Tuozi Cave, Nanjing, China. Beijing: Science Press. 85-130 |
| [24] | Echassoux A, Moigne A M, Moulle P E, et al. 2008. Les faunes de grands mammifères du site de l'homme de Yunxian, Quyuanhekou, Quingqu, Yunxian, Province du Hubei, Republique Populaire de Chine. In: Henry D L, Li T eds. Le Site de l'homme de Yunxian, Quyuanhekou, Quingqu, Yunxian, Province du Hubei. Paris: CNRS Editions. 253-364 |
| [25] | Ellerman J R, Morrison-Scott T C S, 1951. Checklist of Palaearctic and Indian Mammals 1758-1946. London: British Museum (Natural History) Trustees. 1-810 |
| [26] | Faure M, Guérin C, 1983. Le Sus scrofa (Mammalia, Artiodactyla, Suidae) du Gisement Pléistocène Supérieur de Jaurens, a Nespouls, Corrèze, France. Nouv Arch Mus Hist Nat Lyon, 21: 45-63 |
| [27] | Frantz L, Meijaard E, Gongora J, et al. 2016. The evolution of Suidae. Ann Rev Anim Biosci, 4(3): 1-3 |
| [28] | Fu R Y, 2001. The age and characteristics of Quaternary mammalian faunas from northeastern China. In: Deng T, Wang Y eds. Proceedings of the Eighth Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press. 167-176 |
| [29] | Fujita M, Kawamura Y, Murase N, 2000. Middle Pleistocene wild boar remains from NT Cave Niimi Okayama prefecture, west Japan. J Geosci Osaka City Univ, 43: 57-95 |
| [30] | Geraads D, 2004. New skulls of Kolpochoerus phacochoeroides (Suidae: Mammalia) from the Late Pliocene of Ahl al Oughlam, Morocco. Palaeontol Afr, 40: 69-83 |
| [31] | Ghosh M, 1988. The craniology and dentition in the pigmy hog, with a note on the genetic status of Porcula Hodgson, 1847. Rec Zool Surv Ind, 85: 245-266 |
| [32] | Gongora J, Groves C, Meijaard E, 2017. Evolutionary relationships and taxonomy of Suidae and Tayassuidae. In: Melletti M, Meijaard E eds. Ecology, Conservation and Management of Wild Pigs and Peccaries. Cambridge: Cambridge University Press.1-19 |
| [33] | Groves C P, 1981. Ancestors for the pigs:taxonomy and phylogeny of the genus Sus. Technical Bulletin Number 3, Department of Prehistory Research School of Pacific Studies, Australian National University. Canberra: Australian National University Press. 1-96 |
| [34] | Groves C P, 2007. Current views on taxonomy and zoogeography of the genus Sus. In: Albarella U, Dobney K, Ervynck A et al. eds. Pigs and Humans: 10,000 Years of Interaction. Oxford: Oxford University Press. 15-29 |
| [35] | Groves C P, Grubb P, 1993. The Eurasian Suids Sus and Babyrousa. In: Oliver W L R ed. Pigs, Peccaries, and Hippos:Status Survey and Conservation Action Plan. Switzerland: IUCN, the World Conservation Union. 107-111 |
| [36] | Guérin C, Patou-Mathis M ed, 1996. Les grands mammifères Plio-Pléistocènes d'Europe. Paris: Masson. 1-291 |
| [37] | Han D F, 1987. Artiodactyla fossils from Liucheng Gigantopithecus Cave in Guangxi. Mem Inst Vert Paleont Paleoanthrop, Acad Sin, 18: 135-208 |
| [38] | Harris J M, Liu L, 2007. Superfamily Suoidea.In: Prothero D R, Foss S E eds. The Evolution of Artiodactyls. Baltimore: Johns Hopkins University Press. 130-150 |
| [39] | Hooijer D A, 1961. Fossil vertebrates of Ksar Akil, a Palaeolithic rock shelter in the Lebanon. Zool Verh Leiden, 49: 4-65 |
| [40] | Hou S K, 2023. Superfamily Suoidea.In: Zhang Z Q, Hou S K, Wang L H et al. eds. Palaeovertebrata Sinica, Vol III, Fas 9(1): Cetacea and Artiodactyla I. Beijing: Science Press. 93-152 |
| [41] | Hou S K, Deng T, He W, et al. 2014. Foraging behavior of Chleuastochoerus (Suidae, Artiodactyla): a case study of skull and mandible morpho-functional analysis. Sci China-Earth Sci, 57: 988-998, doi: 10.1007/s11430-013-4733-z |
| [42] | Hou S K, Su D F, Kelley J, et al. 2019. New fossil suid specimens from the Terminal Miocene Hominoid Locality of Shuitangba, Zhaotong, Yunnan Province, China. J Mammal Evol, 26: 557-571 |
| [43] | Hu C K, Qi T, 1978. Gongwangling Pleistocene mammalian fauna of Lantian, Shaanxi. Palaeontol Sin, New Ser C, 21: 1-64 |
| [44] | Hu H Q, Yu H D, Wei G B, et al. 2023. The first discovery of Sus xiaozhu skulls from Yanjinggou fauna in Wanzhou, Chongqing. Quat Sci, 43(3): 793-802 |
| [45] | Huang Y P, 1996. Mammalian fossils. In: Tangshan Archaeological Team ed. Locality of the Nanjing Man Fossils 1993-1994. Beijing: Cultural Relics Publishing House. 83-247 |
| [46] | Huang Y P, Fu R Y, Li X, et al. 2021. Studies on the animal fossils from Jinniushan Hominin Site.In: Lü Z E ed. The Jinniushan Hominin Site. Beijing: Cultural Relics Press. 168-285 |
| [47] | Iannucci A, 2024. Sus arvernensis (Suidae, Mammalia) from the early Villafranchian of Villafranca d'Asti (northern Italy): a revision of the largest sample of a key suid species. Boll Soc Paleontol Ital, 63: 45-81 |
| [48] | Iannucci A, Sardella R, Strani F, et al. 2020. Size shifts in late Middle Pleistocene to Early Holocene Sus scrofa (Suidae, Mammalia) from Apulia (southern Italy): ecomorphological adaptations? Hystrix, 31: 10-20 |
| [49] | Iannucci A, Cherin M, Sorbelli L, et al. 2021. Suidae transition at the Miocene-Pliocene boundary: a reassessment of the taxonomy and chronology of Propotamochoerus provincialis. J Mammal Evol, 28: 323-335 |
| [50] | Iannucci A, Pazonyi P, Sebe K, 2024. The Hungarian fossil record of the Pliocene pig Sus arvernensis (Suidae, Mammalia). Swiss J Palaeontol, 143: 39, https://doi.org/10.1186/s13358-024-00335-4 |
| [51] | Jin C Z, Wang Y, Liu J Y, et al. 2021. Late Cenozoic mammalian faunal evolution at the Jinyuan Cave site of Luotuo Hill, Dalian, Northeast China. Quat Int, 577: 15-28 |
| [52] | Kumar S, Gaur R, 2013. First record of maxillary dentition of Potamochoerus theobaldi (Suidae, Mammalia) from the Upper Siwaliks of India. Riv Ital Paleontol Stratigr, 119: 57-63 |
| [53] | Kurtén B, 1968. Pleistocene Mammals of Europe. Chicago: Aldine Publishing Co. 1-317 |
| [54] | Leakey L S B, 1958. Some East African Pleistocene Suidae. Fossil Mamm Afr, 14: 1-132 |
| [55] | Lee Y C, 1963. A new species of Potamochoerus from Chinhsien, Shansi. Vert PalAsiat, 7(2): 161-165 |
| [56] | Leslie D M, Huffman B A Jr, 2015. Potamochoerus porcus. Mamm Spec, 47: 15-31 |
| [57] | Liaoning Museum, Benxi Museum (LMBM), 1986. Miaohoushan—a Site of Early Paleolithic in Benxi County, Liaoning. Beijing: Cultural Relics Publishing House. 1-110 |
| [58] | Liu L P, 2003. Chinese Fossil Suoidea: Systematics, Evolution, and Paleoecology. Helsinki: University of Helsinki. 1-41 |
| [59] | Liu L P, Kostopoulos D S, Fortelius M, 2004. Late Miocene Microstonyx remains (Suidae, Mammalia) from northern China. Geobios, 37: 49-64 |
| [60] | Liu L Q, Bosse M, Megens H-J, et al. 2019. Genomic analysis on pygmy hog reveals extensive interbreeding during wild boar expansion. Nat Commun, 10: 1-9 |
| [61] | Liu W H, Dong W, Zhang L M, et al. 2017. New material of Early Pleistocene Sus (Artiodactyla, Mammalia) from Yangshuizhan in Nihewan Basin, North China. Quat Int, 434: 32-47 |
| [62] | Lü Z E, Huang Y P, Li P S, et al. 1989. Yi Yuan fossil Man. Acta Anthropol Sin, 8(4): 301-313 |
| [63] | Ma A C, Tang H L, 1992. On discovery and significance of a Holocene Ailuropoda-Stegodon fauna from Jinhua, Zhejiang. Vert PalAsiat, 30(4): 295-312 |
| [64] | Martínez-Navarro B, Madurell-Malapeira J, Ros-Montoya S, et al. 2015. The Epivillafranchian and the arrival of pigs into Europe. Quat Int, 389: 131-138 |
| [65] | Martínez-Navarro B, Madurell-Malapeira J, Ros-Montoya S, et al. 2024. The late Villafranchian absence of pigs in Europe. Comment on Iannucci A. The occurrence of suids in the post-Olduvai to pre-Jaramillo Pleistocene of Europe and implications for late Villafranchian biochronology and faunal dynamics. Quaternary 2024, 7, 11. Quaternary, 7: 51 |
| [66] | Mayer J J, Brisbin I L, 1988. Sex identification of Sus scrofa based on canine morphology. J Mammal, 69: 408-412 |
| [67] | Mayer J J, Novak J M, Brisbin I L Jr, 1998. Evaluation of molar size as a basis for distinguishing wild boar from domestic swine:employing the present to decipher the past. In: Ryan K, Quick J eds. Ancestors for the Pigs: Pigs in Prehistory. MASCA Res Pap Sci Archaeol, 15: 39-53 |
| [68] | McKenna M C, Bell S K, 1997. Classification of Mammals above the Species Level. New York: Columbia University Press. 1-631 |
| [69] | Meijaard E, D'Huart J, Oliver W, 2011. Family Suidae (pigs). In: Wilson D E, Mittermeier R eds. Handbook of the Mammals of the World. Barcelona: Lynx Edicions. 248-291 |
| [70] | Neaux D, Blanc B, Ortiz K, et al. 2020. How changes in functional demands associated with captivity affect the skull shape of a wild boar (Sus scrofa). Evol Biol, 48: 27-40 |
| [71] | Orliac M J, Antoine P-O, Ducrocq S, 2010. Phylogenetic relationships of the Suidae (Mammalia, Cetartiodactyla): new insights on the relationships within Suoidea. Zool Scr, 39: 315-330 |
| [72] | Palombo M R, Gallai G, Arca M, et al. 2012. Sus sondaari van der Made, 1999 (Mammalia, Suidae) from Monte Tuttavista (Orosei, western Sardinia,Italy). Riv Ital Paleontol Stratigr, 118(1): 183-191 |
| [73] | Payne S, Bull G, 1988. Components of variation in measurements of pig bones and teeth, and the use of measurements to distinguish wild from domestic pig remains. Archaeozoologia, 2(1): 27-66 |
| [74] | Pearson H S, 1928. Chinese fossil Suidae. Palaeontol Sin, Ser C, 5(5): 1-75 |
| [75] | Pei W C, 1936. On the mammalian remains from locality 3 at Choukoutien. Palaeontol Sin, Ser C, 7(5): 1-108 |
| [76] | Pei W C, Li Y H, 1958. Discovery of a third mandible of Gigantopithecus in Liu-cheng, Kwangsi, South China. Vert PalAsiat, 2(4): 193-200 |
| [77] | Pickford M, 1993. Old World suoid systematics, phylogeny, biogeography and biostratigraphy. Paleontol Evol, 26-27: 237-269 |
| [78] | Pickford M, 2013. Suids from the Pleistocene of Naungkwe Taung, Kayin State, Myanmar. Paleontol Res, 16: 307-317 |
| [79] | Pickford M, 2015. Late Miocene Suidae from Eurasia:the Hippopotamodon and Microstonyx Problem Revisited. München:Verlag Dr.Friedrich Pfeil, 42: 1-126 |
| [80] | Pickford M, Obada T, 2016. Pliocene suids from Musaitu and Dermenji, Moldova: implications for understanding the origin of African Kolpochoerus Van Hoepen E, Van Hoepen H, 1932. Geodiversitas, 38(1): 99-134 |
| [81] | Pilgrim G E, 1926. The fossil Suidae of India. Palaeontol Ind, 8: 1-65 |
| [82] | Qi G Q, Lin Z Y, An J Y, 2006. Animal remains from the Dadiwan site. In: The Gansu Provincial Institute of Cultural Relics and Archaeology ed. Dadiwan in Qin'an—Report on Excavations at a Neolithic Site, Vol II. Beijing: Cultural Relics Publishing House. 861-900 |
| [83] | Qiu Z X, 2000. Nihewan fauna and Q/N boundary in China. Quat Sci, 20: 142-154 |
| [84] | Qiu Z X, 2006. Quaternary environmental changes and evolution of large mammals in North China. Vert PalAsiat, 44(2): 109-132 |
| [85] | Qiu Z X, Qiu Z D, Deng T, et al. 2013. Neogene land mammal stages/ages of China:toward the goal to establish an Asian land mammal stage/age scheme. In: Fortelius M, Wang X M, Flynn L eds. Fossil Mammals of Asia:Neogene Biostratigraphy and Chronology. New York: Columbia University Press. 29-90 |
| [86] | Rook L, Martínez-Navarro B, 2010. Villafranchian: the long story of a Plio-Pleistocene European large mammal biochronologic unit. Quat Int, 219: 134-144 |
| [87] | Ruvinsky A, Rothschild M F, Larson G, et al. 1998. Systematics and evolution of the pig. In: Rothschild M F, Ruvinsky A eds. The Genetics of the Pig. Oxford shire: CABI. 1-16 |
| [88] | Shikama T, 1949. The Kuzuii Ossuaries: geological and palaeontological studies of the limestone fissure deposits, in Kuzuii, Totigi Prefecture. Sci Rep Tohoku Univ, 2nd Ser, 23: 1-209 |
| [89] | Shikama T, Hasegawa Y, 1965. Fossil suid from Kurihama, Kanagawa Prefecture. Sci Rep Yokohama Nat Univ, Sect. 2(12): 37-43 |
| [90] | Smith A T, Xie Y, 2008. A Guide to the Mammals of China. Princeton: Princeton University Press. 1-544 |
| [91] | Souron A, 2012. Histoire évolutive du genre Kolpochoerus (Cetartiodactyla: Suidae) au Plio-Pléistocène en Afrique orientale.Ph. D Thesis. Poitiers: Université de Poitiers. 1-518 |
| [92] | Sun J J, Zhang B, Chen X, et al. 2021. New fossils of Late Pleistocene Sus scrofa from Yangjiawan Cave 2, Jiangxi, China. Vert PalAsiat, 59(1): 64-80 |
| [93] | Suraprasit K, Jaeger J-J, Chaimanee Y, et al. 2016. The Middle Pleistocene vertebrate fauna from Khok Sung (Nakhon Ratchasima, Thailand): biochronological and paleobiogeographical implications. ZooKeys, 613: 1-157 |
| [94] | Suraprasit K, Jaeger J-J, Chaimanee Y, et al. 2021. Taxonomic reassessment of large mammals from the Pleistocene Homo-bearing site of Tham Wiman Nakin (Northeast Thailand): relevance for faunal patterns in mainland Southeast Asia. Quat Int, 603: 90-112 |
| [95] | Tedford R H, Flynn L J, Qiu Z X, et al. 1991. Yushe Basin, China: paleomagnetically calibrated mammalian biostratigraphic standard from the late Neogene of eastern Asia. J Vert Paleont, 11(4): 519-526 |
| [96] | Teilhard de Chardin P, Pei W C, 1941. The fossil mammals from Locality 13 of Choukoutien. Palaeontol Sin, New Ser C, 11: 1-106 |
| [97] | Teilhard de Chardin P, Piveteau J, 1930. Les mammiféres fossils de Nihowan (Chine). Ann Paléontol, 19: 1-134 |
| [98] | Tong H W, Zhang B, Chen X, et al. 2021. Chronological significance of the mammalian fauna from the Early Pleistocene Shanshenmiaozui site in Nihewan Basin, northern China. Acta Anthropol Sin, 40(3): 469-489 |
| [99] | Tong H W, Zhang B, Chen X, 2024. An overview and new insights into the Quaternary mammalian fauna from the Nihewan Basin in North China. Acta Anthropol Sin, 43(1): 157-183 |
| [100] | Van der Made J, 1996. Listriodontinae (Suidae, Mammalia), their evolution, systematics and distribution in time and space. Contrib Tert Quat Geol, 33: 3-254 |
| [101] | Van der Made J, 1999. Biogeography and stratigraphy of the Mio-Pleistocene mammals of Sardinia and the description of some fossils. Deinsea, 7: 337-360 |
| [102] | Van der Made J, 2022. An updated range chart for the Oligocene to recent west Eurasian Suoidea: variation in species richness. Paleontol Evol Mem Esp, 9: 34-36 |
| [103] | Van der Made J, Moya-Sola S, 1989. European Suinae (Artiodactyla) from the Late Miocene onwards. Boll Soc Paleontol Ital, 28: 239-339 |
| [104] | Von den Driesch A, 1976. A guide to the measurement of animal bones from archaeological sites. Peabody Mus Bull, 1: 1-137 |
| [105] | Wang J, 2006. Early Pleistocene mammalian fauna from Bajiazui, Qingyang, Gansu, China and their environment. Master's Thesis. Xi'an: Northwest University. 1-88 |
| [106] | Wu X Z, 2006. Yunxi Man—Excavation Report of the Huanglongdong Site. Beijing: Science Press. 1-271 |
| [107] | Young C C, 1932. On the Artiodactyla from the Sinanthropus site at Chouk'outien. Palaeontol Sin Ser C, 8: 1-158 |
| [108] | Zdansky O, 1928. Die Saugetiere der Quartarfauna von Choukoutien. Palaeontol Sin Ser C, 5: 1-198 |
| [109] | Zeder M A, Lemoine X, 2020. Factors affecting molar size in Sus scrofa. J Archaeol Sci, 124: 105266 |
| [110] | Zhang Y, Zhang S Q, Xu X, et al. 2013. Zooarchaeological perspective on the Broad Spectrum Revolution in the Pleistocene-Holocene transitional period, with evidence from Shuidonggou Locality 12, China. Sci China-Earth Sci, 43: 628-633 |
| [111] | Zhao X T, Gao F Q, Liu C J, 1986. Quaternary stratigraphy and mammalian fossil sites in Qingyang of Gansu. Quat Sci, 7(1): 79-86 |
| [112] | Zheng S H, Han D F, 1993. Mammalian fossils. In: Zhang S S ed. Comprehensive Study on the Jinniushan Paleolithic Site. Mem Inst Vert Paleontol Paleoanthropol, Acad Sin, 19: 43-127 |
| [113] | Zong G F, 1981. Pleistocene mammals from Tunliu, Shanxi. Vert PalAsiat, 19(2): 174-183 |
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