古脊椎动物学报 ›› 2024, Vol. 62 ›› Issue (1): 13-32.DOI: 10.19615/j.cnki.2096-9899.231013
收稿日期:
2023-08-10
出版日期:
2024-01-20
发布日期:
2024-01-17
通讯作者:
* lichun@ivpp.ac.cn基金资助:
WANG Wei1, LEI Hong2, LI Chun1,*()
Received:
2023-08-10
Published:
2024-01-20
Online:
2024-01-17
Contact:
* lichun@ivpp.ac.cn摘要:
现在的中国南方地区在三叠纪时期曾位于特提斯洋东部。在该区域产出的众多海生爬行动物中,恐头龙(Dinocephalosaurus)的形态特化,颈部极度增长,可与时代略晚的长颈龙(Tanystropheus)相比拟。这两种爬行动物分别归属于恐头龙科(Dinocephalosauridae)和长颈龙科(Tanystropheidae), 而这两个科的成员加之另外一些基干物种代表了主龙型类(Archosauromorpha)中一系列早期分异出的支系。报道一件产自云南罗平地区中三叠世安尼期地层的主龙型类成年个体新标本IVPP V18579, 与目前已知的早期主龙型类成员相比,该标本显示出与恐头龙属一致但区别于其他属种的特征,主要包括前颌骨发育有超过外鼻孔后缘的较长后背侧突,方骨前后向较宽大且具有后边缘的凹缺,大部分尾椎上的人字骨带有远端前后向的延展。然而,新标本体型明显小于恐头龙,且显示出与恐头龙属和梳齿龙属(另一恐头龙科成员)不同的鉴别特征,例如吻部窄长、牙齿短锥状且异齿性弱、枢椎和前部颈椎的神经棘显著更高等。基于最新的用于早期主龙型类系统发育分析的特征矩阵,系统发育分析结果确定V18579代表一个新的恐头龙科成员,在恐头龙科中与恐头龙属互为姊妹群关系。于是基于V18579建立了一新属新种:玲珑南蛇龙(Austronaga minuta), 同时对其进行了详细的解剖学特征描述,并讨论了它可能具有的水生适应性。
中图分类号:
王维, 雷洪, 李淳. 云南中三叠世一恐头龙科小型主龙型类. 古脊椎动物学报, 2024, 62(1): 13-32.
WANG Wei, LEI Hong, LI Chun. A small-sized dinocephalosaurid archosauromorph from the Middle Triassic of Yunnan, southwestern China. Vertebrata Palasiatica, 2024, 62(1): 13-32.
Fig. 1 Holotype of Austronaga minuta (IVPP V18579) discovered from Luoping, Yunnan, China, with the skull, anterior cervical vertebrae, and most of the caudal vertebrae preserved on a block
Fig. 2 Skull and anterior cervical vertebrae (A) with interpretative line drawings (B, C) of the holotype of Austronaga minuta (IVPP V18579) from the Middle Triassic in Luoping, Yunnan, China Abbreviations: an. angular; ar. articular; atic. atlas intercentrum; atpl. atlas pleurocentrum; atn. atlas neural arch; ax. axis (pleurocentrum); axic. axis intercentrum; axn. axis neural spine; bo. basioccipital; cop. coronoid process; cr(3-5). (third to fifth) cervical rib; crat. cervical rib of atlas; crax. cervical rib of axis; cv(3-6). (third to sixth) cervical vertebrae (centra); cv(3-5)n. (third to fifth) cervical vertebral neural arches; d. dentary; ept. ectopterygoid; f. frontal; ipv. interpterygoid vacuity; j. jugal; la. lacrimal; m. maxilla; mte. maxillary teeth; n. nasal; p. parietal; pbs. parabasisphenoid; pfor. parietal foramen; pl. palatine; pm. premaxilla; po. postorbital; prf. prefrontal; pt. pterygoid; q. quadrate; rap. retroarticular process; so. supraoccipital; sq. squamosal; sran. surangular
Item | Length | Item | Length (centrum) | Height at midpoint | |
---|---|---|---|---|---|
Skull | 47.5 | Atlas | - | - | |
Snout | 24.1 | Axis | 3.1* | 6.4 | |
Orbit approximately length | 15.9 | Cervical V3 | 6.1 | 5.6 | |
Temporal region | 7.5 | Cervical V4 | 9.0 | 5.1 | |
Mandible | 46.6 | Cervical V5 | 10.4 | 5.0 |
Table 1 Measurements of the holotype of Austronaga minuta (IVPP V18579) from the Middle Triassic in Luoping, Yunnan, China (mm)
Item | Length | Item | Length (centrum) | Height at midpoint | |
---|---|---|---|---|---|
Skull | 47.5 | Atlas | - | - | |
Snout | 24.1 | Axis | 3.1* | 6.4 | |
Orbit approximately length | 15.9 | Cervical V3 | 6.1 | 5.6 | |
Temporal region | 7.5 | Cervical V4 | 9.0 | 5.1 | |
Mandible | 46.6 | Cervical V5 | 10.4 | 5.0 |
Fig. 3 Middle region of the tail (A) with interpretative line drawings (B) of representative caudal vertebrae in the holotype of Austronaga minuta (IVPP V18579) from the Middle Triassic in Luoping, Yunnan, China Abbreviations: cav. caudal vertebra (centrum); cavn. caudal vertebral neural arch; ch. chevron
Fig. 4 Phylogenetic results showing the position of Austronaga minuta in Dinocephalosauridae and other relationships within Archosauromorpha A. strict consensus tree; B. majority-rule reduced consensus trees(nodes with bootstrap proportion > 50% remained with values labeled above)
char. 1-50 | 11?30?1111 | ??[0/1]0000??? | ?0?0000-01 | 11002?0000 | ???0?00001 |
char. 51-100 | ???00????? | ??????0?1? | ?0?1??0??? | ?????????1 | 0?????-??1 |
char. 101-150 | -------?1? | ?????????? | ?????????? | ?????????? | ???000?1?0 |
char. 151-200 | ?01?10?11? | 01100001[0/4]0 | 00010?1220 | 21??210211 | 1?00????2[1/2] |
char. 201-250 | ?????????? | ?????????? | ?03??????? | ?????????? | ?????????? |
Appendix Scores of character states for the holotype of Austronaga minuta
char. 1-50 | 11?30?1111 | ??[0/1]0000??? | ?0?0000-01 | 11002?0000 | ???0?00001 |
char. 51-100 | ???00????? | ??????0?1? | ?0?1??0??? | ?????????1 | 0?????-??1 |
char. 101-150 | -------?1? | ?????????? | ?????????? | ?????????? | ???000?1?0 |
char. 151-200 | ?01?10?11? | 01100001[0/4]0 | 00010?1220 | 21??210211 | 1?00????2[1/2] |
char. 201-250 | ?????????? | ?????????? | ?03??????? | ?????????? | ?????????? |
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