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    15 September 2008, Volume 46 Issue 3
    AN EARLY DEVONIAN (PRAGIAN) SARCOPTERYGIAN FROM ZHAOTONG, YUNNAN, CHINA
    LU Jing, ZHU Min
    2008, 46(3):  161-170. 
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    A new basal dipnomorph sarcopterygian, Arquatichthys porosus gen. et sp. nov., from the Lower Devonian (Pragian) of northeastern Yunnan, China, is described on the basis of a nearly complete mandible and several isolated scales. It closely resembles basal dipnomorphs in following characters: short and L-shaped pit-line of infradentary 4 (uniquely shared with dipnomorphs), small parasymphysial dental plate attachment, three coronoids, broad “tooth pavement” on the lateral portion of coronoids, large adductor fossa, irregularly scattered pores of mandibular sensory canal on the lateral surface of mandible, and distinct anteroventral process of rhomboid scale. Arquatichthys has a unique mandible morphology characterized by a strongly arched upper margin, and an elongated overlapped area by quadratojugal and maxillary. As the second dipnomorph from the Pragian of China, Arquatichthys adds evidences to understanding the early diversification of sarcopterygians. The character transitions of scales are discussed within the framework of the interrelationships of early sarcopterygians.
    A NEW ARMORED PLACODONT FROM THE MIDDLE TRIASSIC OF YUNNAN PROVINCE, SOUTHWESTERN CHINA
    ZHAO Li-Jun, LI Chun, LIU Jun, HE Tao
    2008, 46(3):  171-177. 
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    A new cyamodontoid placodont, Glyphoderma kangi gen. et sp. nov., is established on the basis of an articulated skeleton from the Middle Triassic Zhuganpo Member of the Falang Formation, Yunnan, southwestern China. It differs from the geologically younger Psephochelys polyosteoderma (early Late Triassic) in having a more complicated carapace osteoderm structure, and 3 enlarged tubercular osteoderms fused to the posterior surface of the skull. Glyphoderma is the only Ladinian member of the family Placochelyidae, all other placochelyids being Late Triassic. Placodonts, including both armored and non-armored forms are distributed from the Anisian to the Carnian in China.
    A NEW GENUS AND SPECIES OF CAUDIPTERID DINOSAUR FROM THE LOWER CRETACEOUS JIUFOTANG FORMATION OF WESTERN LIAONING, CHINA
    HE Tao, WANG Xiao-Lin, ZHOU Zhong-He
    2008, 46(3):  178-189. 
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    A new oviraptorosaur, Similicaudipteryx yixianensis gen. et sp. nov. is described from the Jiufotang Formation (120 Ma) of the Jehol Group in western Liaoning, China, which is referred to the Caudipteridae based on a dagger-like pygostyle and the shape of the ilium that are most similar to those of Caudipteryx. It differs from other oviraptorosaurids in that the ratio of pubis to ilium length is 1.46 and the presence of two large and deep hypapophyses on dorsal vertebrae. The known caudipterids have previously been found only from the Jianshangou Member of the Yixian Formation (125 Ma) of the Sihetun area in Liaoning Province. S. yixianensis represents the first caudipterid dinosaur from the Jiufotang Formation. The new discovery provides more information for the discussion of the evolution of oviraptorids during the Early Cretaceous and adds to the dinosaur assemblage of the Jehol Biota.
    A NEW SPECIES OF THE LATE MIOCENE TAPIRS (PERISSODACTYLA, TAPIRIDAE) FROM THE LINXIA BASIN IN GANSU, CHINA
    DENG Tao, HE Wen, CHEN Shan-Qin
    2008, 46(3):  190-209. 
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    The tapirid fossils from the Neogene deposits are relatively scarce, mainly including isolated teeth. In Europe, some Miocene specimens of the genus Tapirus were found, among which,the earliest species was Tapirus telleri established by Hofmann (1893) from Goriach in Austria, and Opole 2 in Poland with the age of Astaracian(MN 6). Another widespread species was Tapirus priscus established by Kaup(1833) from Eppelsheim, Westhofen, and Wissberg in Ger-many, Can Llobateres in Spain, Priay in France, Biedermannsdorf in Austria, and Rudabanya in Hungary, with the age of Vallesian (MN 9)(Heissig,1999). Zdansky (1935) reported the Late Miocene Tapirus leilhardi from Wuxiang, Shanxi, but its locality and age were inexact.Recently, the anterior portion of a juvenile skull and mandible(IVPP V 15523) was found from the upper red clays of the Liushu Formation at Shilei(IVPP LX 0031) in Guanghe, Linxia, and then the anterior portion of an adult skull(V 15522) was found from the same horizon at Wangjiashan(LX 0501) in Hezheng, Linxia. The discovery fills the gap and vacuum in evolution of Tapiridae in East Asia, because the origin of the Quatemary tapirs in China has been uncertain. IVPP LX and IVPP V are the locality and specimen prefixes of Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, respectively. Terminology of lower cheek teeth is according to Qiu and Wang(2007, fig.7) for perissodactyls.
    PLIOCENE BIOSTRATIGRAPHIC SEQUENCE IN THE NIHEWAN BASIN, HEBEI, CHINA
    LI Qiang, ZHENG Shao-Hua, CAI Bao-Quan
    2008, 46(3):  210-232. 
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    A sequence of fluvio-lacustrine and paludine deposits with thickness over 150 meters occurs in the Nihewan Basin, Hebei, China(Yuan et al.,1996; Min et al.,2006).A series of ~30m(="90 feet") thick red homogeneous clay exposured near the Sanggan River Gorge (=Shixia) was originally considered as the weathering matter of the base rocks(Barbour et al.,1926). Due to the absence of fossils, its age once was speculated to belong to the Pontian (Teilhard and Piveteau,1930) or the Late Miocene Baodean(Black et al.,1933). However, a sequence of ~30 m thick red gravel-bearing clay at the bottom of the section of Dahonggou in Shixia was named as"Dahonggou Formation"by Chen(1988). At the same time, the~12 m and~7 m thick red clays in the lower part of the Hongya Nangou and Pump Station, respectively, were also included into this Pliocene formation.A series of 20~30 m thick redish or yellow-redish alternating sandy clay and gravel beds are exposed in the Luanshigedagou near Hongya village. Yielding Hipparion and Chilotherium, this deposit was considered to belong to the"Hipparion Red Clay"with a Pliocene age(Huang et al.,1974). The sequence of~45 m thick red clay containing sandy gravel and lens of calcareous nodules in the Huabaogou near Xiyaozitou village was divided into the upper"Yuxian Formation"and the lower"Huliuhe Formation"by the composition of large fossil mammals. These two formations were respectively correlated to be equal in age to the Pliocene Jingle Formation and the Late Miocene Bahe Formation(Wang,1982). In view of the illegibility of the boundary and the lack of significant differences in the fossil faunas between these two formations, Zhang et al.(2003) incorporated them both in the Late Pliocene Yuxian Formation.A series of fluvio-lacustrine and paludine deposits exposed on the both sides of the Huliu River were named "Daodi Formation"and considered of Late Pliocene age(Du et al.,1988). The original sections used for erecting the"Daodi Fm." are located in the Laowogou near Daodi, the Nangou near Hongya, the Jiangjungou near Xiyaozitou, the Hougou near Qijiazhuang, the Yuanzigou near Yuanzi, the Xiaoshuigou near Qianjiashawa, the Lianjiegou near Beimajuan, the Niutoushan(="Pulu") near Pulu and the Danangou near Dongyaozitou. Zhang et al.(2003) figured that these short sections all belong to the"Yuxian Fm.", so they suggested that"Daodi Fm."should be abolished and attributed to the"Yuxian Fm.", whereas Cai et al.(2004) retained both"Daodi Fm."and "Yuxian Fm.". The red clay stratum from the first layer at the bottom of the Laowogou section is stillunnamed due to the uniformity of the grain size, the absence of gravels and aquatic animal remains. Based on the limited fossil mammals, the age of this stratum was thought to be the late Middle or early Late Pliocene(Zhang et al.,2003; Cai et al.,2004). The Pliocene/Pleistocene boundaries, on the other hand, have already been established in the Laowogou, Donggou, Taiergou and Niutoushan sequences (Cai et al.,2004; Zheng et al.,2006; Min et al.,2006; Cai et al.,2007). Judging from the known information, the lower part of the exposed strata in the Nihewan Basin should be attributed to the Late Pliocene Yuxian Formation or some unnamed red clays.These outcrops exist in the Sanggan River Gorge area and on the both sides of the lower reaches of the Huliu River with lithology of aeolian clay, fluvio-lacustrine red clay contained gravels and paludine sandy clay. On the both sides of the lower reaches of the Huliu River, there are 12 localities and sections, in which one or more layers of fossil mammals have been found(Fig.1). The purpose of this paper is to update a faunal list of these mammals based on the review of specimens and their localities, and also an attempt to correlate the other sections to the Laowogou section by comparing their mammalian compositions and lithological strata. Finally,a Late Pliocene biostratigraphic sequence and the environmental changes in the Nihewan Basin are discussed.
    EARLY PLEISTOCENE SUID (MAMMAL) FROM THE DAJUSHAN, HUAINAN, ANHUI PROVINCE (CHINA)
    DONG Wei 
    2008, 46(3):  233-246. 
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    A karstic cave called Laodong with rich Late Miocene mammals was found in 1990 in the Dajushan(117°01′E,3235′N) area,15 km northwest of Huainan City, Anhui Province in eastern China(Jin et al.,2000; Jin,2004). During the implementation of the State Key Project of the 9h Five-year-plan", Jin and colleagues found two other fossil localities. One of them is located in a fissure called Xiliexi(it means West Fissure in Chinese) and about 60 m west of the Laodong Cave. Many mammal fossils were unearthed from the fissure deposits. Some specimens of Sus lydekkeri were identified as below.